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In the future, mankind has been left to a planet called Zion. They have been forced on to this planet by space creatures called "Victims." Terrorized by these Victims, Zion remains the only human inhabited planet left, thus mankind's last hope for survival. But with the alien invasion of Victims moving in, their only defense are the five "Goddesses" and their pilots. This brings in Zero: headstrong and short-tempered. Zero hopes to one day fulfill his dream of becoming a goddess pilot.

Centuries ago, mankind abandoned planet Earth after the Vice, a race of mechanical aliens, drove them from their home into a life adrift in space. Now resigned to starships forged of pieces of Earth's shattered moon, the final hope for humanity lies in the hands of Actresses, young women born with the ability to wield the only weapons that can harm the Vice: Alice Gears, mechanical suits that can finally turn the tide against the alien incursion.